Lists of the insect types in the Zoological Museum, University of
Helsinki – Ephemeroptera
Hans Silfverberg
Based on:
Silfverberg, H. 1989: Lists of the insect types in the Zoological
Museum, University of Helsinki. 10. Ephemeroptera. - Acta Entomol.
Fennica 55:1-2.
The types of 8 taxa in the species group within Ephemeroptera are
listed. References to original descriptions are given and, when
the taxon is presently known under a different name, to an explanatory
work. Additional information is given concerning J. A. Aro, and
the loss of some of his types.
This paper lists 7 species and 1 subspecies of Ephemeroptera, the
types of which are deposited in the Helsinki Museum. Unless otherwise
indicated, the specimens are dry, pinned adults. Some of the species
were described by Johan Emil Aro (1874-1928), a school teacher in
Viipuri/ Viborg (today Vyborg in Russia), who described his species
in Finnish; in his earlier work (Aro 1910) he did not mention anything
about the material he had used. It has been possible to find from
later works (Aro 1928, Tiensuu 1939) information concerning possible
type specimens, but it was decided to restrict this category to specimens
that Aro himself has collected. In the Helsinki Museum there are older
materials, which Aro at least had seen insomuch as be listed his species
from such provinces from which he had not himself collected; none
of them has any label in Aro's handwriting.
Aro was killed in a traffic accident on 5.2.1928 (before his later
work was finished), and no information could be found concerning the
fate of his material. It seems either to have been disposed of immediately,
or if preserved in the school where he had worked, or in the Viipuri
Natural History Museum, then it was undoubtedly destroyed during the
Second World War. For two species thus no type material seems to exist.
However, it seems useful to give some information concerning these
two species, too, especially since Aro's papers are not easily available.
Baetidae
saliens Tiensuu 1939:112; Baetis. Syntype from Russia
(Karelian Rep.): Sortavala, Tiensuu, 2.9.34. Listed as synonym of
Baetis subalpinus Bengtsson, 1917 (Müller-Liebenau 1969).
Heptageniidae
convergens Aro 1910:31; Echdyurus. 3 syntypes (in
alcohol) from Finland: Pori, joen varrelta [by the river], 24.V.03,
Aro; same except 1.VI.03; Liperi, pappilan ranta [vicarage shore],
16.VI.05. Listed as synonym of Heptagenia fuscogrisea (Retzius,
1783) (Tiensuu 1939).
mirabilis Aro 1910:30; Cinygma. 27 syntypes (all in
alcohol) from Finland: Vihti, lätäkköjä Kirjavan
rannassa [ponds on the shore of Kirjava], 21/5 1903, J. E. Aro (6
larvae in one tube, 2 in another); Kokemäki, joelta [by the river],
29.VII.04, Aro (1 larva); Sysmä, maantieltä Nuoramoisjärven
kohdalla [on the road at Lake Nuoramoisjärvi], 14.VI.06, Aro
(1 male); Kuopio, Tuovilanlahti, 6.VII.01, J. E. Aro (1 male, 1 larva);
Kuopio, Tuovilanlahti, 2.VII.01, J. E. Aro (2 larvae); Tuovilanlahti,
joesta puitten alta [from the river under trees], 8/VII 01, Aro (9
larvae); Tuovilanlahti, 14/VII 01 (4 larvae). Listed as synonym of
Arthroplea congener Bengtsson, 1908 (Tiensuu 1939).
Ephemerellidae
brunnescens Tiensuu 1935:16; Melanameletus. Holotype
from Russia (Karelian Rep): Sortavala, Tiensuu, 18.VI.1931. Listed
as synonym of Eurylophella karelica Tiensuu, 1935 (Puthz 1978).
karelica Tiensuu 1935:21; Eurylophella. 10 syntypes
(nymphs, in alcohol) from Russia (Karelian Rep.): Kurkijoki, kirkon
kylän läpi virtaavasta savisesta ojasta [from a clayey ditch
running through the church village], 30/7 1934. Leg. L. Tiensuu.
Polymitarcidae
ladogensis Tiensuu 1935:8; Polymitarcys. 61 syntypes
from Russia (Karelian Rep.): Salmi, Tiensuu, 1776, Type 6539 (1 ex.,
dry); Salmi, Tiensuu, Lunkulansaari, 11.8.1934 (1 ex., dry); Salmi,
Tiensuu, Lunkulansaari, 12.8.1934 (1 ex., dry); Salmi, Tiensuu, Lunkulansaari,
13.8.1934 (1 ex., dry); Salmi, Tiensuu, 14.8.1934 (2 exx., dry); Salmi,
Tiensuu (15 exx, dry); Salmi, Karkku, 16.8.1934, leg. L. Tiensuu,
Type 6538 (15 exx. in alcohol in one tube); Salmi, Lunkulansaari,
2.8.1934, leg. L. Tiensuu, Type 6540 (4 larvae and 2 adults in alcohol
in one tube); Salmi, Tiensuu (3 larvae, 7 subimagines and 5 adults
in alcohol in one tube); Kl: Salmi, leg. L. Tiensuu, prep. no 1 (1
ex., in alcohol and some parts on slide); Kl: Salmi, leg. L. Tiensuu,
prep. no 2 (same as the former). Tiensuu wrote: "Types 6538-6540 in
Mus.Zool.Helsinki" but gave no additional information on the type
specimens, and therefore I consider his entire original material syntypic.
Listed as synonym of Ephoron nigridorsum (Tshernova, 1934)
(Illies 1967).
Caenidae
dimidiata var. fennica Aro 1928:29; Caenis.
- see fennica.
fennica Aro 1928:29; Caenis dimidiata var. 6 syntypes
from Russia (Leningrad obl.): Jääski, J. E. Aro; Antrea,
Kuorekoski, J. E. Aro (3 exx., numbered 377, 557 and 952); Muola,
Hotakka, J. E. Aro, 24.VII.1921; Muola, Äyräpää,
J. E. Aro, 29.VII.1921. (Jääski is nowadays known as Lesogorskij,
Antrea as Kamennogorsk and Muolaa as Kirillovskoe.) Listed as simple
synonym of Caenis horaria (Linnaeus, 1758) (Saaristo 1966).
undosa Tiensuu 1939:122; Caenis. 6 syntypes from Russia
(Karelian Rep.): Salmi, Tiensuu, Lunkulans., 13.8.34 (1 ex.) and Salmi,
Tiensuu (5 exx.). Listed as synonym of Caenis nocturna Bengtsson,
1917 (Saaristo 1966), which again nowadays is considered a synonym
of Caenis lactea (Burmeister, 1839) (Puthz 1978).
Types not found
Siphlonuridae
fennica Aro 1910:3 1; Palmenia. Described from a few
specimens collected in Finland near Oulu on 20.5.1906 and 25.5.1906.
Types apparently lost. Listed as synonym of Parameletus chelifer
Bengtsson, 1908 (Tiensuu 1939).
Heptageniidae
flavomaculatus Aro 1928:54; Ecdyurus. Described from
three specimens collected in Finland at Sysmä: Suopelto in 6.8.1902.
Types apparently lost. Listed as synonym of Ecdyonurus joernensis
Bengtsson, 1909 (Illies 1967), which is nowadays known as Nixe
joernensis (Bengtsson) (Flowers 1986).
References
- Aro, J. E. 1910: Piirteitä päiväkorennoisten (Ephemeridae)
elämäntavoista ja kehityksestä. - Viipurin suomalaisen
realilyseon vuosikertomus, p. 1-32.
- Aro, J. E. 1928: Suomen päivänkorennoiset (Ephemerida).
- Otavan hyönteiskirjasia 3 / Vanamon kirjoja 27:1-68.
- Flowers, R. W. 1986: Holarctic distribution of three taxa of
Heptageniidae (Ephemeroptera). - Entomol. News 97:193-197.
- Illies, J. 1967: Ephemeroptera. - In Illies (ed.), Limnofauna
Europaea. Stuttgart, pp. 212-219.
- Müller-Liebenau, I. 1969: Revision der europäischen
Arten der Gattung Baetis Leach, 1815 (Insecta, Ephemeroptera). -
Gewässer und Abwässer 48/49:1-214.
- Puthz, V. 1978: Ephemeroptera. - In Illies (ed.), Limnofauna
Europaea. 2. edition. Stuttgart, pp. 256-263.
- Saaristo, M. 1966: Revision of the Finnish species of the genus
Caenis Steph. (Ephemeroptera). - Ann. Entomol. Fennici 32:68-87.
- Tiensuu, L. 1935: On the Ephemeroptera-fauna of Laatokan Karjala
(Karelia Ladogensis). - Ann. Entomol. Fennici 1:3-23.
- Tiensuu, L. 1939: A Survey of the Distribution of Mayflics (Ephemerida)
in Finland. - Ann. Entomol. Fennici 5:97-124.
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