Notes on Hierochloë australis

Hierochloë australis (Schrad.) Roem. & Schult., three morphs from Finland: the common morph with brownish violet tinge in the panicle (Lohja, Osuniemi, 1986; on the left), a morph with green panicle (Lohja, Osuniemi, 1986; in the middle), and a morph with acuminate glumes (Lammi, Untulanharju, 1994; on the right). Image: Harri Harmaja (scanned from dried specimens). The above brownish and green plants, collected quite near-by, were at a rather early stage of development, before the anthesis. H. australis (Poaceae) of Finland mostly has brownish tinge to its inflorescence, plants with green panicles being rather infrequent. These two morphs also display some additional, minor differences; e.g., the anthers are pale brownish in the brown morph but yellow in the green morph. Some measurements of the pollen grains showed them to be ca. 28-43 µm in diametre in the species, those of the green morph tending to be slightly smaller on the average. However, it remains to be studied whether the above differentiation, detected in a few populations, is traceable in a wider material.
    The right-hand plants in the image are truly deviating from the bulk of Finnish H. australis through their acuminate (vs. short-acute) glumes. Additional differences include a more contracted panicle and more acute lemmas as well as later flowering (unless the morph is sterile?).
    One more observation: the (brownish-panicled) plants of a stand (Finland, Lammi, Jahkola, 1998) possessed a distinct additional smell besides the normal coumarine scent. A rather strong, disagreeable smell resembling that of Stachys sylvatica (Lamiaceae) was present in fresh plants (due to the activities of some endophytic fungus?).

Created September 13, 2004.